Grounded J171, pin 3 to chassis ground. No change.
I'm still a bit worried about the crackling sound on power-up (well, the no audio problem is also an issue, of course). It seems to correspond to the flickering VUs. Once the machine has stabilised, the lights are at full brightness and the crackling sound stops. It also seems to correspond to what I assume would be the delay time before the relays are supposed to kick in (or out). There must be some correlation between the crackling and strange VU meter behaviour and the non-tripping relays because, if I'm understanding this correctly, they are on the same rail.