Great tune...I'll cut to the chase.
I'm gonna' go out on my own little limb here and say that the guitar tone in the chorus is actually *PERFECT* for this song if the bass can become more a part of the song. It's a real mid-heavy git tone, and it will sit very well, but IMHO, the bass in this song is mixed way too low. I think, as is, this needs to be mixed more like a rock tune...heavier on the bass and high end.
I'd try turning the bass up a few db; everything else is recorded so cleanly, I really don't think mud will be a problem.
Oh, and I feel your pain on the bass tuning thing. I've learned to use the tuner on the guitars, and ditch it on the bass...use your ears and monitor really loudly when you're playing. If it's off, you'll know it immediately. Everybody knows that no matter what the tuning meter says, it can be off for some mysterious reason with the rest of the song.
Bass sounds in tune to me, though...I can't be sure because IT'S MIXED TOO LOW...blah blah blah...you get my point.
Great collab....Tastey stuff, Gidge. Great song, dres.