I’m new to commenting on mixing so this is probably all about my taste or lack of it, rather than anything technical. Or it could just be my headphones?
I find that the rhythm guitars are panned too far apart leaving the centre of the field very open – exposing the bass, lead and vocal to exist in there own space. This maybe what you want? I know I tend to like big wall of sound arrangements.
I think that is part of the bass issues, without even a 'bleed' of guitar from left or right it seems separate from the other guitarists and I think that is why you feel they are out of beat; when I don't think they are.
The bass I would love to hear fatter you’ve got all those ‘high grit’ harmonics in the other guitars I’d love to hear some really deep stuff going on with the lower end – I think the tonality of your vocals would sit even better with that thickness behind them.
I find the guitars all very similar in tone, even the lead – what does it sound like in a mono mix? Can you distinguish all the elements?
Your vocals are so distinct (I have always enjoyed them when you’ve posted on song writing), and the band’s musicianship excellent. The song itself is engaging and has a great feel.
HTH
Burt