Ok, it appears to my unaided eye that you are using the main outs from the mackie into the 1010. Is that correct? If so, this is not the cleanest signal path and there are so many ways to adjust level that my axons are hurting.
I would suggest using either the direct outs or the channel inserts. See this thread about it:
Mackie - 1010 connection thread
And you will need to connect a cable from each Mackie channel to a 1010 input channel. If you use the insert jack from the Mackie, then level setting will be controlled by the channel trim and 1010 control panel. If using the direct (almost) on the Mackie then the trim AND the fader will adjust the level.
Now the big question is: What are the settings for the 1010 control panel inputs? Select Monitor tab and then adjust the Mixer inputs.
The -4.1db to -2.5 peaks are plenty hot enough. Maybe what is messing with your mind is the fact that the Mackie is running in the analog domain and can output signals beyond 0 dB, but once that pristine audio enters the virtual universe it can never go beyond digital 0. Every signal must live in a sub-zero environment. Hence that cold digital sound.

So your ultimate level reference needs to come from the 1010 control panel, assuming you haven't overloaded the previous Mackie gain stages which can result in that harsh analog sound.
I think you'll find if you connect using the direct outs or insert jacks and setting the 1010 input levels from the control panel that setting levels will take less axons and decibels. Now, back to the music. Hope this helps,
- Jim